• Professor
  • Department Chair, Interim
  • Ecology and Integrative Biology
  • Ph.D.
  • Department of Biology
  • Temple University

I am an ecologist who studies planktonic organisms in marine and fresh waters. My research emphasis is on protists (algae and protozoa) and their trophic interactions. A continuing thread in my research is the physiology and ecological importance of mixotrophic protists that are both photosynthetic and phagotrophic. See my website for recent publications and teaching.